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View of Shore grykes along the western Istrian coast

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SHORE GRYKES ALONG THE wESTERN ISTRIAN COAST ALONG THE wESTERN ISTRIAN COAST OBALNI GRAJKI OB ZAHODNI ISTRSKI OBALI

Stefano FURLANI1, 3, Davide CHERSICLA2, Guido BRESSAN2, Sara BIOLCHI3, 4 & Franco CUCCHI3

Izvleček UDK 551.4(497.5-16)

Stefano Furlani, Davide Chersicla, Guido Bressan, Sara Biol- chi & Franco Cucchi: Obalni grajki ob zahodni Istrski obali V članku prestavimo nove topografske in morfološke izsledke ter fizikalno-kemične parametre izmerjene na izbrani� obal- ni� grajki� severoza�odne obale Istre med Savudrijo in Zambratijo. Raziskave smo omejili na šest profilov s štirimi do petimi plimskimi bazeni. V vse� profili� smo določili tri tipične morfološke cone, kateri� značilnosti so tesno pove- zane z lokalnim plimovanjem. Višji bazeni so izpostavljeni za- krasevanju, zato so v nji� oblike bolj gladke, v nji�ovem dnu pa prevladujejo sedimenti s kopnega, predvsem terra rossa. V smeri morja opazimo vse večji vpliv biokorozije. Ob vznožju grajkov najdemo peske in prodnike. Kemični parametri kažejo, da so nižje ležeči grajki izpostavljeni predvsem delovanju mor- ske vode, medtem ko višje ležeče oblikuje predvsem deževnica.

Razvoj obalni� grajkov je povezan z nezveznostmi, ki so po- sledica lokalni� tektonski� deformacij.

Ključne besede: obalni kras, za�odna Istra, obalni grajki, plimski bazeni, apnenec.

1 Dipartimento di Geografia, Università degli Studi di Padova, Italy, e-mail: sfurlani73@gmail.com

2 Dipartimento di Scienze della Vita, Università degli Studi di Trieste, Italy

3 Dipartimento di Geoscienze, Università degli Studi di Trieste, Italy

4 Dipartimento di Scienze della Terra, Università di Modena e Reggio Emilia, Italy Received/Prejeto: 9.9.2010

Abstract UDC 551.4(497.5-16)

Stefano Furlani, Davide Chersicla, Guido Bressan, Sara Biol- chi & Franco Cucchi: Shore grykes along the western Istrian coast

we provided new data on topograp�y, morp�ology and p�ysi- cal/c�emical parameters (pH, T, NONO2-, Ca2+, PO43-, NaCl) col-) col- lected in several s�ore grykes along t�e Nort�western Istrian coast, between Savudrija and Zambratija. Six transects, eac�

containing four to five pools, �ave been surveyed. Three mor- p�ological zones �ave been identified along t�e selected pro- files. Morp�ological features of t�e s�ore grykes along t�e western Istrian coast are, in fact, closely related to t�e local tide. Hig�-level pools are affected by karstic processes, and t�e surface is usually smoot�. At t�eir bottom, terrigenous depos- its, mainly terra rossa, occur. Seaward, bioerosion prevails and at t�e bottom of t�e grykes, sand and rounded pebbles �ave been found. C�emical/p�ysical parameters suggest t�at grykes located at lower altitudes are affected by seawater factors, w�ile pools located at increasing altitudes are affected mainly by rainfall and consequentially fres�water or saltwater remaining from rainfalls or storm events. S�ore gryke genesis is strongly controlled by geological weakness, along w�ic� t�ey develop.

Their origin is in fact due to local tectonics, w�ile t�eir develop- ment is related to t�e active vertical tectonic subsidence of t�e study area. Pools located at �ig�er altitudes are mainly affected by solution karst processes, but due to t�e tectonic downdrop of t�e area, w�en t�e grykes come in contact wit� sea, t�ey are gradually s�aped by marine processes.

Keywords: coastal karst, western Istria, s�ore grykes, tide- pools, limestone.

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The morp�ology of limestone coasts is determined by t�e presence of karst morp�ologies w�ic� �ave been in�er- ited as a result of c�anges in t�e relative sea level, and consequently modified by marine processes (Tren�aile 1987). This mec�anism is well-pronounced along recent- ly subsiding areas, suc� as t�e Eastern Adriatic, w�ere t�e development of coastal morp�ologies is conditioned by vertical tectonic movements.

In t�e intertidal zone along western Istria, t�anks to t�e gently sloping coast and t�e outcrop of limestone rocks, grykes, c�annels and, to a small extent, tidepools occur. Grykes are solutionally enlarged vertical or steep- ly inclined joints in t�e surface of a karstland, extending for up to a few meters into t�e limestone (Monroe 1970).

A gryke is composed by several pools, sometimes co- alescent or joined t�roug� sip�ons and filled by seawater during �ig� tides or storms.

w�ile tidepools on limestone coasts �ave been ex- tensively studied (Ley 1979; Denny & Gaines 2007), no relevant works can be found regarding s�ore grykes and c�annels. Alt�oug� some considerations and met�ods regarding t�e study of tidepools can be exported to t�e study of s�ore grykes, t�e peculiarity and complexity of t�ese morp�ologies suggest use of different s�ape index- es. Regarding t�e study of tidepools, Feldmann (1937), distinguis�ed temporary pools and permanent pools depending on t�e time period in w�ic� t�ere is water.

These coastal features are, in fact, prominent structures are, in fact, prominent structures t�at form in t�e intertidal zone w�ere resistant bedrock is exposed (Griggs 2007). The type of rock exposed, t�e

tidal range, t�e wave action, t�e erosion and t�e weat�er- ing processes acting on t�e rocks are significant in t�eir genesis and development (De waele et al. 2009).

The intertidal belt is affected by variable c�emical/

p�ysical conditions. It is subjected to seasonality, cir- cadian differences and to rainfall, w�ic� can dilute t�e seawater, making it brackis�. Hig� biodiversity adds in- terest to t�e studied morp�ologies. Denny and Gaines (2007) recently provided a complete overview on tide- pools. They are �abitats of adaptable fauna and flora t�at, because of �uge waves, strong currents and exposure to midday sun, fluctuations in sea level, water temperature, salinity and oxygen content, must be able to cope wit� a c�anging environment.

Few studies concern t�e morp�ology of limestone coasts along t�e Eastern Adriatic (D’Ambrosi 1948; Forti 1985; Perica et al. 2004; Fouac�e et al. 2000; Antonioli et al. 2007; Furlani et al. 2009), and no papers �ave dealt specifically wit� t�e fact t�at s�ore morp�ologies deepen

�ere, despite t�e favourable conditions for t�eir genesis and evolution. The present work started from a t�esis in environmental sciences (C�ersicla 2009), w�ic� provid- ed data to study t�e Istrian limestone s�ore grykes.

The paper aims to provide new data on topograp�y, morp�ological zonation and p�ysical/c�emical param- eters (pH, T, etc.) collected on several s�ore grykes along t�e nort�western Istrian coast, along six transects at Sa- vudrija and Zambratija, and to discuss t�e evolution of t�e studied morp�ologies.

INTRODUCTION

STUDY AREA

GEOLOGICAL SETTING

The study area is located between Savudrija and Zam- bratija (Nort�western Croatia, Fig. 1). In t�is sector of t�e Istrian coast, Early Cretaceous limestones, in par- ticular Albian and Cenomanian in age, crop out. They correspond to a period of t�e evolution of t�e Adriatic Carbonate Platform (Vla�ović et al. 2005) during w�ic�

it was affected by periods of emergences due to t�e inter- action of tectonics and eustatic c�anges. Lower Aptian deposits, in particular, represent an event connected wit�

t�e partial drowning of t�e platform. During t�e transi- tion between Aptian and Albian, t�e regional emersion of t�e platform occurred. Albian deposits are usually t�in-bedded and are c�aracterised by typical marginal features (desiccation cracks, ripple marks and dinosaur

tracks). So during t�e Barremian, Aptian and Albian most of t�e platform was dominated by s�allow subtidal environments ric� in Foraminifera, wit� t�e gradual rise of rudists. During Late Cretaceous t�e platform attained its maturity and began to disintegrate (Velić et al. 2002).

This process was accompanied by almost continuous synsedimentary tectonic activity, eustatic c�anges and extensive development of rudist communities. Lower Cenomanian deposits are c�aracterised by facies affected by t�e synsedimentary tectonic defomation of t�e plat- form. After Middle Cenomanian a gradual re-unification occurred, wit� s�allow-marine deposition (Vla�ović et al. 2005).

The area is c�aracterised by a gentle tectonic setting.

The main structural framework is t�e Buje Anticline,

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w�ic� stretc�es from Savudrija, across Buje, Oprtalj and Buzet, following t�e wNw-ESE direction. Considering t�e extension (45 km) of t�e structure, its genesis �ad to be followed by deformation of wider areas (Matičec 1994). Considering t�at t�e Dinaric structures differ in extension, orientation and strike from t�ose parallel to t�e Buje Anticline, t�e last one s�ould �ave been formed during a neotectonic p�ase, younger t�an Oligocene and Miocene, as a consequence of a recent global stress (NNE-SSw, Matičec 1994).

Seismotectonic activity is not relevant, even if t�e Nort�weastern Adriatic s�ores are acknowledged as be- ing c�aracterized by recent submersion. The underwa- ter position of a number of intertidal morp�ologies or coastal ant�ropic structures (notc�es, platforms, Roman age remains, etc.) supports t�is idea (Fouac�e et al. 2000;

Antonioli et al. 2004, 2007; Auriemma & Karinja 2008;

Furlani et al. 2010a, b; Faivre et al. 2010) and suggests t�at, in t�e Nort�western Istrian area, 2 ka BP t�e sea level was -1.6 m below msl, indicating vertical tectonic rates up to -0.77 mm/yr.

ENVIRONMENTAL SETTING

Rainy winters and �ot dry summers c�aracterise t�e Mediterranean climate in t�e Istrian area. Mean an- nual rainfall measured in Trieste averages 1015 mm/yr (Stravisi 2003). The minimum mean value was recorded in February, w�ereas t�e daily peak (105 mm) normally occurs in November. Storms are more frequent at t�e end of t�e summer and during autumn. In western Istria t�e mean rainfall is lower (Rovinj site, 766 mm/yr).

On average, t�ere are 130 days/year w�en precipita- tion occurs. In a year t�ere are approximately 31.5% rainy days, 64.4% sunny days, 3.0% snowy days and 1.1% days wit� �ail (Tommasini 1979). The average annual temper- ature along t�e studied coast is approximately 16°C. The

�ottest mont� is August (24°C), w�ereas t�e coldest one is January, �aving temperatures lower t�an 6°C.

The area is c�aracterised by a prevalence of winds blowing from t�e first quadrant, mainly from t�e ENE (Bora) (Carrera et al. 1995). Sout�-easterly winds (Sci- rocco or Jugo) are important because of t�e wide fetc�

(over 800 km). Tides are semi-diurnal, wit� mean Fig. 1: Location of the transects in the study area. The base map (a) was created using SRTm data (Jarvis et al. 2008), while the geologi- cal map (c), that includes the location of the transects, is redrawn from Plenicar et al. (1973). b) The wind rose shows the dominant winds in the area (Stravisi 2003).

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spring-tide values of 0.86 m in Trieste, and mean neap- tide values of 0.22 m (Dorigo 1965; Polli 1970). The con- comitance of spring tides, seic�es, sout�-easterly winds and low atmosp�eric pressure can cause a sea-level rise of 1.60 m. Along t�e nort�ern Istrian coast, tides are 80% of t�ose at Trieste. The winds of t�e Sout�west and t�e Nort�west quadrant raise it, t�us leading to “�ig�- water” p�enomena. Mean significant wave �eig�t during t�e year is lower t�an 0.5 m (Dal Cin & Simeoni 1994), w�ile t�e �ig�est offs�ore wave �eig�t, bot� for Bora and Scirocco storms, is about 5 m (Cavaleri et al. 1996).

The �ig�est mean �ourly speed for sout�east wind in t�e period 1958-1987 was 27.3 m/s.

An estimate of wave energy for t�e Nort�ern Adri- atic area in t�e form of H2T2 (H and T corresponding to t�e significant wave �eig�t and period) was studied using t�e complete series of 2000-2003 t�ree-�ourly measurements available from t�e RON-APAT (Fontolan et al. 2007) wave gauge located offs�ore Ancona (latitude 43.3704600, longitude 13.3001300). The annual value is 15.61 m2s2.

Mean sea temperatures range between 9 and 11°C, wit� peaks in August (24°C). Seawater salinity displays

�ig�er values in winter t�an during summer and ranges between 34 and 39 psu (Miramare oceanograp�ic buoy).

Frost is rare and affects mainly t�e s�eltered areas of t�e bays.

Biogeoc�emical parameters, on a seasonal-clima- tological basis, in t�e Nort�ern Adriatic Sea �ave been studied by Zavatarelli et al. (1998). Surface NONO2- distri- bution during winter decreases in t�e wE direction, so it is moved by barocline circulation. Moreover it is lower during spring-summer, probably because it is used by plants for metabolic processes. POPO43- is usually very low in t�e nort�ern Adriatic (Degobbis 1990; Krom et al.

1991; Giani et al. 2003), and it decreases in t�e eastward direction, so t�e nort�ern Adriatic Sea can be consid- ered p�osp�o-limited. O2 concentrations are �ig�er t�an 8 mg/l, sometimes �ig�er t�an 14 mg/l. Maxi- mum values �ave been observed during spring because of low temperatures and algal activity, w�ile minimum values �ave been observed during summer, mainly be- cause of increasing temperatures and salinity. Seawater in t�e Gulf of Trieste is generally �ig�er t�an 8 mg/l O2 (www.arpa.fvg.it).

MATERIALS AND METHODS

TOPOGRAPHICAL MEASUREMENTS Six perpendicular s�ore transects (Figs. 2 & 3) �ave been measured using t�e met�od suggested by Hunt (2007) and Antonioli et al. (2007) to measure coastal mor- p�ologies and Roman remains. Figures 2 and 3 represent sc�ematic transects; in fact only t�e t�res�old and t�e bottom of t�e gryke �ave been reported, considering t�e particular importance of t�ese parameters. The bottom is, in fact, generally flat w�ile t�e t�res�old is t�e “gate”

t�roug� w�ic� t�e water flows into t�e gryke.

Altitudinal measurements and distances �ave been collected using, respectively, an optical (Salmoirag�i Ertel automatic level), or a laser level and a measur- ing tape. we measured (Table 1) t�e elevation (m msl) of t�e bottom of t�e gryke, t�e �eig�t of t�e gryke (m), t�e distance from t�e s�ore (m), t�e lengt� (L, m), t�e widt� (l, m) and t�e eccentricity (L/l), obtained as t�e ratio between lengt� and widt�. we use t�e terms used by Lewis (1964), t�at is: �ig�-level pools (or grykes) to indicate t�e pools located at �ig�er altitudes above sea level (supratidal zone), and mid-level (intertidal) or low- level (subtidal) to t�e ot�ers.

Altitudinal measurement consists of: 1) measure- ment of t�e dept� of t�e marker wit� respect to t�e lo-

cal sea level at t�e time of surveying. Values reported in Table 1 represent t�e mean value of multiple measures;

2) correction of surveyed measures wit� respect to t�e nearest tide gauge data collected at t�e time of survey- ing. Error bars for t�e elevation and age values of t�e sea level markers �ave been provided in order to consider t�e inaccuracy of measures.

Elevation measurements were collected via optical (Salmoirag�i Ertel automatic level) or mec�anical in- struments (invar rod) or wit� a laser level. The survey- ing was performed during periods of low wave energy in order to minimize t�e errors. Measurements �ave been reduced to t�e mean sea level applying tidal corrections at t�e surveyed sites, using t�e data of t�e Trieste tide gauge. In fact, tide amplitude is particularly significant in t�e nort�ern Adriatic Sea, as it can be up to ~2 m. El- evation measurements are given (Stravisi & Purga 2005) wit� respect to t�e Italian reference plane network of t�e Istituto Geografico Militare (Genova Mean Sea Level 1942; Gamboni 1965).

Limestone samples �ave been collected using a geo- logical �ammer. The rock identification �as been carried out t�roug� t�in sections in t�e Department of Geosci-

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ences in Trieste. The geograp�ic locations �ave been ob- The geograp�ic locations �ave been ob- tained using a GPS Garmin 12®.

The multivariate analysis was carried out using SPSS software, by means of topograp�ical variables (bottom dept� elevation and distance from t�e s�ore). The Eucli- dean quadratic distance �as been used as index, and t�e ward’s criterion was used for t�e cluster-analysis.

CHEMICAL/PHYSICAL MEASUREMENTS Salinity (psu). NO2-, Ca2+, PO43-, NaCl and pH �ave been collected using field instruments during winter, spring and summer, 2008 (Tab. 2-4). C�emical/p�ysical data

�ave been reported in Tab. 2-4 depending on t�e season.

The day and �our of measurement are reported for every Fig. 2: Topographical transects at Savudrija. The thick line indicates the mean sea level. Capital letters indicate the pools on the surveyed grykes. A is the gryke at the highest elevation, while the following letters indicate as lower grykes.

Fig. 3: Topographical transects at zambratija. The thick line indicates the mean sea level. Capital letters indicate the pools on the surveyed grykes. A is the gryke at the highest elevation, while the following letters indicate as lower grykes.

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transect. “No data” indicates t�at it was not possible to measure because t�e pool was empty.

The Tetratest® kit, a fast and effective qualitative met�od, was used to measure dissolved O2 and nutrients (NO2-, NO3-, Ca2+, PO43-); salinity was measured using t�e Hand Refractometer S/mill-E ATAGO® w�ile t�e temperature, pH and salinity were collected using t�e multiparametric waterproof Hand�eld pH/Cond Meter xS PC650.

SELECTION OF TRANSECTS

Six transects, marked by roman numbers (I to VI), per- pendicular to t�e s�ore and located between t�e lower tidal zone and t�e supratidal zone �ave been studied. They are composed of a number of pools, grykes, breac�es and c�annels named wit� capital letters (A is t�e morp�ology located most inland, and t�e letters progress toward t�e s�ore, Figs. 2 & 3).

Four transects are located near t�e Lig�t�ouse of Savudrija (Figs. 1 & 2), w�ile two transects are located near Zambratija (Figs. 1 & 3).

They �ave been c�osen wit� respect to t�e follow- ing criteria:

• t�e proximity between t�e transects;

• all t�e transects are roug�ly E-w oriented, so wind and wave action is similar;

• every transect �as different slope degrees and dif- ferent elevations above mean sea level;

• t�e dimension and s�ape of t�e studied morp�ol- ogies are different in every transect;

• in Savudrija t�e transects are lig�tly affected by ant�ropic influence due to t�e presence of a camping area;

• in Zambratija t�e study area is not affected by �u- man presence.

RESULTS

Collected data are reported in Tab. 1-5. In Tab. 1 topo- grap�ical and morp�ological data are reported; in Tab. 2, 3 and 4 p�ysical/c�emical parameters collected respec- tively during winter (Tab. 2), spring (Tab. 3) and summer (Tab. 4), w�ile in Tab. 5 t�e results of t�e cluster analysis are reported.

SITE DESCRIPTION

The topograp�ical and morp�ological description of t�e pools is reported in Tab. 1. The surveyed pool, t�e eleva- tion of t�e bottom (m msl), t�e �eig�t of t�e pool (m), t�e minimum distance from t�e s�ore (m), t�e eccentric- ity (ratio between lengt� and widt�), t�e nature of de- posits on t�e bottom of t�e pool and t�e nature of slope weat�ering �ave been considered.

The location of t�e transects at Savudrija (Fig. 5) and t�e number of morp�ologies surveyed is reported following:

•  transect I (45°29’23”N; 13°29’26”E): it is located just below t�e Savudrija Lig�t�ouse. It consists of five pools (Fig. 2);

• transect II (45°29’22”N; 13°29’26”E): it is located a few meters sout� of t�e first transect. It consists of four pools (Fig. 2);

• transect III (45°29’20”N; 13°29’27”E): it is located 70 m sout� from transect II. It consists of four coalescent pools (Fig. 2);

• transect IV (45°29’18”N; 13°29’27”E): it is located a few meters sout� of transect III, near an artificial ce-

mented pool. It can be considered a meandered c�annel;

letters indicate significant points, mainly in correspon- dence to t�e curves of t�e meander (Fig. 2);

Fig 4: Dendrogram of the hierarchical classification of the shore grykes considering the topographical data.

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The location of t�e transects at Zambratija (Fig. 6) and t�e number of morp�ologies surveyed is reported following:

• transect V (45°29’18”N; 13°29’27”E): it is located 1500 m sout� of transect IV. It consists of four pools (Fig. 3);

• transect VI (45°29’18”N; 13°29’27”E): it is located a few meters sout� of transect V. It consists of five pools (Fig. 3).

TOPOGRAPHICAL RESULTS

Topograp�ical results �ave been reported in Tab. 1. The el- evation (m msl) of t�e bottom of t�e studied grykes rang- es between -0.75 m (transect IV) and 1.35 m (transect I).

The �eig�t of t�e t�res�old of t�e grykes ranges between 1.47 m (transect I) and -0.59 m (transect V). The maxi- mum distance from t�e s�ore is 18.5 m (transect IV). The eccentricity ranges between 6.25 and 0.18. In many cases it was impossible to calculate eccentricity because of t�e complex s�ape of t�e gryke. Deposits on t�e bottom of t�e grykes are mainly terrigenous (terra rossa) in t�e

�ig�-level grykes (Fig. 7), w�ile grykes at lower altitudes are filled wit� sand or rounded pebbles. The bottom of one pool (IC) is cemented. Observations on t�e rock surface suggest t�at biocorroded surfaces are located at lower altitudes, w�ile at �ig�er altitudes karstic solution prevails. Some surfaces at �ig�er altitudes seem to be smoot�ed by running waters.

Tab. 1: Topographical and geomorphological data of the surveyed morphologies. The table reports: the name of the transect and number of the surveyed pool (pool is generically intended as erosional feature), the elevation of the bottom of the pool (m msl), the maximum height of the pool (m msl), the mean distance of the pool from the shore (m); the eccentricity, that is the ratio between the length and the width, the nature of the deposits accumulated at the bottom of the pool and the nature of slope weathering.

Topography Morphology

Pool Bottom

elevation (m) Total height (m)

Distance from the shore

(m)

Eccentricity

(L/l) Deposits on the bottom Slope weathering

I A 1.35 1.47 10.5 2 terrigenous smooth

I B 0.8 0.98 8.5 4.25 terrigenous smooth

I C 0.51 0.67 1 n.p. man-made karstic solution

I D -0.18 0.07 -2 0.36 rounded clasts biocorroded

I E -0.31 0.21 -4.8 0.22 rounded clasts biocorroded

II A 0.36 0.55 7.6 2 terrigenous karstic solution

II B 0.31 0.63 4.2 4.12 angular clasts karstic solution

II C -0.02 0.35 0.6 1.85 angular clasts biocorroded

II D -0.16 -0.06 -0.2 0.18 rounded clasts biocorroded

II E -0.53 -0.29 -1.8 0.48 rounded clasts biocorroded

III A 0.71 1.00 4.3 1.71 terrigenous smooth

III B 0.32 0.67 2.2 1.36 terrigenous karstic solution

III C 0.12 0.34 0.3 3 small clasts biocorroded

III D -0.21 -0.10 -1.5 1.67 rounded clasts biocorroded

IV A 0.52 0.72 18.5 n.p. terrigenous karstic solution

IV B 0.39 0.59 15.5 n.p. terrigenous karstic solution

IV C 0.16 0.36 10.5 n.p. small clasts karstic solution

IV D 0.02 0.22 8.5 n.p. small clasts karstic solution

IV E -0.01 0.66 4 n.p. small clasts karstic solution

IV F -0.43 0.33 -5 1.57 rounded clasts biocorroded

IV G -0.75 -0.01 -6.5 2.5 rounded clasts biocorroded

V A 0.51 0.57 11.5 5 terrigenous karstic solution

V B 0.12 0.18 6.7 2.67 terrigenous karstic solution

V C -0.46 -0.21 -8.8 5.5 sand and clasts biocorroded

V D -0.54 -0.59 -12.5 6.25 sand and clasts biocorroded

VI A 0.35 0.45 14 2.08 terrigenous karstic solution

VI B -0.23 -0.08 2 2 sand and clasts biocorroded

VI C -0.12 -0.02 -4 5 rounded clasts biocorroded

VI D -0.52 -0.34 -5 2 rounded clasts biocorroded

VI E n.p. -0.52 -5.5 n.p. rounded clasts biocorroded

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Fig. 5: Shore grykes at Sa- vudrija: a, b) view of pools at Savudrija collected re- spectively during low and high tide; c) notch mor- phologies bordering pools of the transect Iv; d) pools at higher altitudes of the transect I (Photos: S. Fur- lani).

The multivariate analysis (Tab. 5, Fig. 4) �ig�lig�ts a strong positive correlation between t�e clusters; t�e con- sidered topograp�ical parameters decrease proportion- ally between an element of cluster A and an element of cluster M or B (Tab. 5). Pools belonging to cluster A can be roug�ly identified wit� t�e pools of t�e spray zone (supralittoral), w�ile pools belonging to cluster M and B are t�e grykes located in t�e intertidal or subtidal zone.

LITHOLOGICAL DESCRIPTION

Savudrija transects (Fig. 2): four limestone beds, Albian in age, occur. The bedding strike direction is w 335 N wit� respective dip angle of 10. From t�e bottom to t�e top, beds are defined as follows:

• 60 cm laminated dark grey limestone wit� spo- radic subrounded millimetric or centimetric w�ite plas- ticlasts c�aracterised by wackestone texture;

• 20 cm grey limestone wit� several millimetric or centimetric w�ite plasticlasts wit� wackestone texture;

• 60 cm fossiliferous lig�t grey limestone wit�

grainstone texture, c�aracterised by several Rudist frag- ments;

Fig. 6: Shore grykes at zambratija: a) the fractures have been shaped mainly by mechanical abrasion; b) view of the transect vI. At the base of the pools, terra rossa occurs (Photos: S. Fur- lani).

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Tab. 2: Physical/chemical parameters collected during winter.

pool Temp.

(°C) O₂

(mg/l) NO₂

(mg/l) NO₃

(mg/l) Ca²

(mg/l) PO₄³¯

(mg/l) pH Salinity

(psu) date and hour

I A 9.0 14 < 0.3 10 120 0 9.68 0

04/02 10.00

I B 8.9 14 < 0.3 12.5 120 0 9.37 5

I C 9.5 14 < 0.3 0 580 0 8.91 50

I D 9.8 14 < 0.3 0 580 0 8.12 38

I E 10.5 14 < 0.3 0 580 0 8.24 38

II A 9.0 11 < 0.3 12.5 625 0 8.49 25

13/03 10.30

II B 8.7 11 < 0.3 12.5 650 0 8.35 29

II C 9.5 11 < 0.3 12.5 725 0 8.22 37

II D 9.6 14 < 0.3 12.5 500 0 8.28 36

III A 8.5 8 < 0.3 12.5 650 0 7.60 11

13/03 12.00

III B 9.3 11 < 0.3 12.5 675 0 8.59 25

III C 9.7 14 < 0.3 12.5 700 0 8.29 36

III D 9.7 14 < 0.3 12.5 700 0 8.35 35

IV A 6.5 8 < 0.3 12.5 900 0 8.56 46

13/03 13.00

IV B 3.9 8 < 0.3 12.5 900 0 8.49 65

IV C 2.9 11 < 0.3 12.5 675 0 8.70 46

IV D 5.0 11 < 0.3 12.5 675 0 no data 43

IV E 6.1 11 < 0.3 12.5 675 0 7.92 44

IV F 6.1 14 < 0.3 12.5 500 0 8.35 41

IV G 8.7 14 < 0.3 12.5 500 0 8.36 35

V A 8.0 11 0.3 0 440 0 8.18 38

07/02 13.00

V B 13.3 11 < 0.3 0 540 0 8.62 39

V C 9.2 11 < 0.3 12.5 540 0 8.15 38

V D 10.2 11 < 0.3 12.5 540 0 8.17 37

VI A 12.8 8 < 0.3 0 480 0 8.22 27

07/02 13.00

VI B 10.1 11 < 0.3 12.5 580 0 8.31 36

VI C 9.1 11 < 0.3 12.5 580 0 8.16 37

VI D 9.4 11 < 0.3 12.5 580 0 8.16 40

VI E 10.1 11 < 0.3 12.5 580 0 8.21 40

Tab. 3: Physical/chemical parameters collected during spring.Physical/chemical parameters collected during spring.

pool Temp.

(°C) O₂

(mg/l) NO₂ (mg/l) NO₃

(mg/l) Ca²

(mg/l) PO₄³¯

(mg/l) pH Salinity

(psu) date and hour

I A no data no data no data no data no data no data no data no data

09/06 12.30

I B no data no data no data no data no data no data no data no data

I C 25.7 11 < 0.3 0 625 0 8.60 36

I D 23.1 14 < 0.3 0 550 0 8.42 39

I E 22.0 14 < 0.3 0 500 0 8.27 39

II A 28.5 8 < 0.3 12.5 500 0 8.33 20

09/06 12.00

II B 28.1 11 < 0.3 12.5 500 0 8.46 26

II C 24.7 14 < 0.3 0 500 0 8.18 35

II D 23.9 14 < 0.3 0 500 0 8.27 36

III A 19.8 8 < 0.3 0 650 0 8.07 13

09/06 11.30

III B 26.7 14 < 0.3 0 675 0 8.94 24

III C 24.0 14 < 0.3 0 550 0 8.60 40

III D 25.4 14 < 0.3 0 500 0 8.19 34

IV A 27.0 8 0.8 12.5 700 0 9.07 2

09/06 10.30

IV B 25.2 8 0.3 12.5 675 0 9.06 4

IV C 28.5 11 0.3 12.5 600 0 8.66 28

IV D 26.3 11 < 0.3 12.5 600 0 8.66 30

IV E 26.3 14 < 0.3 0 520 0 8.66 30

IV F 25.3 14 < 0.3 0 550 0 8.12 36

IV G 22.0 14 < 0.3 0 550 0 8.27 38

V A no data no data no data no data no data no data no data no data

09/06 13.30

V B 23.2 14 < 0.3 0 480 0 8.31 38

V C 21.0 14 < 0.3 0 480 0 8.30 40

V D 21.0 14 < 0.3 0 580 0 8.30 40

VI A 23.5 11 < 0.3 0 440 0 8.29 39

09/06 13.00

VI B 23.7 8 < 0.3 12.5 480 0 8.35 38

VI C 22.8 14 < 0.3 0 580 0 8.34 39

VI D 22.6 14 < 0.3 0 580 0 8.30 39

VI E 21.7 14 < 0.3 0 580 0 8.32 40

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Tab. 5: mean values of variables in the clusters.mean values of variables in the clusters.

cluster Bottom height

(m) Elevation

(m m.s.l.)

Distance from the shore

(m)

A -0.20 0.20 1.86

M -0.29 -0.11 -5.28

B -0.65 -0.45 -12.74

Tab. 4: Physical/chemical parameters collected during summer.Physical/chemical parameters collected during summer.

pool Temp.

(°C) O₂

(mg/l) NO₂

(mg/l) NO₃

(mg/l) Ca²

(mg/l) PO₄³¯

(mg/l) pH Salinity

(psu) date and hour

I A no data no data no data no data no data no data no data no data

11/09 14.30

I B no data no data no data no data no data no data no data no data

I C 29.4 14 < 0.3 8 no data 0 8.95 35

I D 26.2 11 < 0.3 0 no data 0 8.49 35

I E 27.0 11 < 0.3 0 no data 0 8.18 35

II A 30.5 14 < 0.3 12.5 no data 0 8.61 37

03/09 16.00

II B 29.0 11 < 0.3 12.5 no data 0 8.67 32

II C 27.9 14 < 0.3 0 no data 0 8.40 35

II D 26.5 14 < 0.3 0 no data 0 8.26 34

III A no data no data no data no data no data no data no data 94

03/09 14.30

III B 27.1 14 < 0.3 0 560 0 8.24 31

III C 26.3 14 < 0.3 0 500 0 8.04 35

III D 26.6 14 < 0.3 0 500 0 8.32 35

IV A 22.0 8 0.8 12.5 160 0.5 8.00 0

03/09 10.30

IV B 24.3 11 0.3 25 140 0.3 7.93 4

IV C 23.5 11 0.5 12.5 240 0.2 7.94 17

IV D 23.9 8 < 0.3 12 380 0 8.05 20

IV E 25.0 11 < 0.3 8 440 0 8.09 18

IV F 25.6 14 < 0.3 0 520 0 8.19 36

IV G 25.6 14 < 0.3 0 520 0 8.19 36

V A 29.4 11 0.3 0 no data 0 8.07 33

11/09 12.30

V B 27.3 14 < 0.3 0 no data 0 8.48 32

V C 26.2 14 < 0.3 0 no data 0 8.35 34

V D 26.2 14 < 0.3 0 no data 0 8.35 34

VI A 24.7 11 < 0.3 0 no data 0 8.09 35

11/09 10.00

VI B 25.1 8 < 0.3 0 no data 0 8.16 34

VI C 25.0 11 < 0.3 0 no data 0 8.20 35

VI D 25.4 11 < 0.3 0 no data 0 8.20 35

VI E 25.4 11 < 0.3 0 no data 0 8.20 35

• 3–5 cm laminated grey limestone wit� sporadic Rudist fragments.

Zambratija transects (Fig. 3): t�ree limestone beds, Cenomanian in age, occur. The bedding strike direction is E 120 S wit� respective dip angle of 12. From t�e bot- tom to t�e top, beds are defined as follows:

• 60 cm lig�t grey limestone wit� mudstone-wacke- stone texture;

• 40 cm grey very fossiliferous limestone wit� pack- stone texture; t�ere are several Rudists, intact or in frag- ments;

• 60 cm lig�t grey limestone wit� mudstone tex- ture.

Fig. 7: a, b) Terra rossa and karst deposits in the pools at higher altitudes at Savudrija (Photos: S. Furlani).

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CHEMICAL/PHYSICAL PARAMETERS Collected data are reported in Tab. 2-4, depending on t�e season. Sometimes, t�e absence of water prevented t�e measurements.

During winter (Tab. 2), temperatures ranged be- tween 2.9 and 13.3°C, O2 ranged between 8 and 14 mg/l, NO2 was 0.3 mg/l or smaller, NO3 ranged between 0 and 12.5 mg/l, Ca2+ ranged between 120 and 900 mg/l, PO4³- was always 0, pH ranged between 7.60 and 9.6, w�ile sa- linity ranged between 0 and 65 psu.

During spring (Tab. 3), temperatures ranged be- tween 19.8 and 28.5°C, O2 ranged between 8 and

14 mg/l, NO2 was smaller t�an 0.3 mg/l up to 0.8 mg/l, NO3 ranged between 0 and 12.5 mg/l, Ca2+ ranged be- tween 480 and 700 mg/l, PO4³- was always 0, pH ranged between 8.07 and 9.07, w�ile salinity ranged between 2 and 40 psu.

During summer (Tab. 4), temperatures ranged between 22.0 and 30.5°C, O2 ranged between 8 and 14 mg/l, NO2 was smaller t�an 0.3 mg/l up to 0.5, NO3 ranged between 0 and 12.5 mg/l, Ca2+ ranged between 140 and 560 mg/l, PO4³- ranged between 0 and 0.5 mg/l, pH ranged between 7.93 and 8.95, w�ile salinity ranged between 0 and 94 psu.

DISCUSSION

The western Istrian area is of interest for t�e occurrence of s�ore grykes. Few tidepools �ave been recognized, prob- ably due to t�e structural and lit�ological conditions and t�e active vertical downdrop. Tectonic rates indicate t�at t�e area �as been subsiding at least since Roman times; in fact arc�aeological remains lie below sea level (Antonioli et al. 2004, 2007). The pools along t�e grykes are usually coalescent, resulting in an elongated and continuous en- larged fracture. The elongated form of t�e studied coastal features and t�eir joint-controlled origin suggested use of t�e term s�ore gryke.

The topograp�ical surveying �ig�lig�ts t�at t�e transects collected at Savudrija are s�orter t�an t�ose collected at Zambratija, because t�e coast is flatter in t�e latter location and t�is allows t�e development of wider coastal platforms. However, t�e longest transect (IV) is located at Zambratija. Its meandering s�ape could be due to t�e coalescence of perpendicular joint sets enlarged by subaerial or marine processes, depending on t�e alti- tude. The measurements of t�e elevation of t�e seaward t�res�old of t�e pools allow determination of t�e eleva- tion at w�ic� t�e single pool comes in contact wit� t�e sea and it is filled by seawater. At Savudrija, t�e elevation of t�e �ig�-level pools is �ig�er t�an at Zambratija, be- cause of t�e morp�ological setting of t�e coast.

The multivariate analysis of topograp�ical param- eters suggests t�e occurrence of 3 clusters, marked by t�e letters A, M and B, w�ic� are strongly related to t�e alti- tude and t�e distance from t�e s�ore (Tab. 5). Ley (1979), studying t�e tidepools along t�e Bristol c�annel, recog- nized a twofold division of t�e fores�ore, and �e divided t�e karren above and around mid-tide and t�e karren located in t�e lower fores�ore. The latter are considered residual forms. The dendrogram (Fig. 4) �ig�lig�ts a strong positive correlation between t�e clusters; t�e con-

sidered topograp�ical parameters decrease proportion- ally between cluster A and cluster M or B (Tab. 5). Pools belonging to cluster A can be roug�ly identified wit� t�e pools of t�e spray zone (supralittoral), w�ile pools be- longing to cluster M and B are t�e grykes located in t�e intertidal or subtidal zone. Generally, differences in alti- tude correspond to differences in emersion time, w�ic�

result in extreme c�anges in p�ysical conditions over a few vertical meters (Harley 2007). As s�own in Tab. 1, t�e pools at �ig�er altitudes (cluster A) �ave a smoot�ig�er altitudes (cluster A) �ave a smoot�

surface; at t�eir bottom, sub-rounded or rounded peb- bles prevail. Locally, terra rossa �ave been observed also seaward w�ere, generally, bioerosion increases surface roug�ness. The cluster M includes mainly t�e pools lo- cated around mean sea level.

The p�ysical conditions along t�e grykes vary greatly, but in general t�e larger and deeper a pool and t�e lower its position on t�e s�ore t�e more it will cor-its position on t�e s�ore t�e more it will cor- respond to a sublittoral �abitat, as suggested by Lewis (1964) in terms of temperature, salinity etc. Hig�-level pools can be significantly different from data reported for t�e Adriatic sea (Giani et al. 2003). However, often t�e pools are coalescent, so t�e topograp�y prevents t�e differentiation of t�e single pool. Variances can be meas- ured mainly during low tides, in pools t�at remain iso- lated from t�e sea, even for several days or weeks. Dur- ing winter, t�e pools IV B and IV C and during spring t�e pool III A s�owed lower temperatures t�an seawater, mainly because t�ey are far from t�e s�ore, wit�out con- tact wit� t�e sea. Hig�er temperatures �ave been found in t�e same pools during summer. Differences can be ascribed mainly to t�e strong evaporation. Even �ig�

salinity values are due to t�e same process (e.g. 94 psu in t�e pool IIIA during summer). Lower values, w�ic�

usually are associated wit� low values in Ca2+ (I A and

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I B during winter), suggest a rainfall input. Mid-level and low-level pools, w�ic� �ave a continuous salt water return from t�e sea, �ave more stable values similar to seawater. Here in fact prevails bioerosion. Low values of PO43- or NO2- �ave been recorded w�ile O2 s�ows values similar to t�e sea. Transect IV �ig�lig�t t�e most sig- nificant differences, mainly during spring and summer, probably due to t�e presence of a waste pipe from t�e camping at t�e back. As a consequence, NO3-, NO2- and PO43- was particularly �ig�, No morp�ologies related to

�uman activity �ave been recognized in correspondence wit� t�is transect.

The collected data suggest t�e occurrence of a tidal zonation of t�e processes t�at influence t�e evolution of t�e s�ore grykes, but t�e genesis seems to be strong- ly related to structural and lit�ological conditions of t�e study area. In fact, t�e studied grykes always occur In fact, t�e studied grykes always occur along geological weaknesses and usually s�ow elongated forms. Even if t�e biological distribution �as not been explored deeply in t�is work, Torunski (1979) suggested t�at erosion rates along t�e Nort�eastern Adriatic coast increase in t�e intertidal zone due to bioerosion. Cucc�i et al. (2006) and Furlani et al. (2009) instead suggested t�at t�e combined action of marine weat�ering process- es leads to an increase in t�e lowering rates up to about

10 times. The close relations between inland and marine grykes force us to relate t�eir origin to an active verti- cal tectonic subsidence, recognized by several aut�ors (Fouac�e et al. 2000; Antonioli et al. 2004, 2007) along t�e western Istrian coast. Tectonic rates indicate t�at t�e area �as been subsiding at least since Roman times (-0.77 mm/y), reinforcing marine processes on karstic features.

The parameters t�at control t�e evolution of t�e grykes are t�e same as t�ose t�at control t�e formation of coast- al karren already studied by Perica et al. (2004): t�e coast inclination in relation to bedding, t�e lit�ological nature of t�e bedrock and t�e occurrence of joints and t�e axis of t�e folds. Thanks to t�e tectonic downdrop of t�e area, grykes are gradually s�aped by marine processes, mainly bioerosion and sometimes by mec�anical abra- sion, w�ere rounded pebbles prevail.

Regarding t�e processes, corrosion acts on t�e lime- stones following t�e variations in c�emical and p�ysical parameters occurring at t�e pools at different elevations.

Consequently, �ig�-level pools are influenced mainly by karstic solution and t�ey are often smoot�ed. Mid- and low-level pools are instead influenced by marine weat�- ering, mainly biological, w�ic� increases t�e roug�ness of t�e limestone surface.

CONCLUSIONS

Morp�ological features of t�e s�ore grykes along t�e western Istrian coast are closely related to t�e local tide.

Hig�-level pools are influenced by karstic processes and t�e surface is usually smoot�. At t�eir bottom, terrige- nous deposits, mainly terra rossa, occur. Seaward, bioero- sion prevails and at t�e bottom of t�e grykes, sand and rounded pebbles �ave been found. However, t�e c�emi- cal/p�ysical parameters suggest t�at grykes at lower alti- tudes are affected by seawater factors, w�ile pools located at increasing altitudes are gradually affected by rainfall and fres�water or saltwater remaining from t�e occur- rence of rainfalls and storm events.

S�ore grykes genesis is strongly controlled by joints, fractures and t�e axis of folds; usually t�ey s�ow elongat- ed forms, along t�e geological weakness. Their origin is due to local tectonics, w�ile t�eir development is related to t�e active vertical tectonic subsidence of t�e study area. Pools located at �ig�er altitudes are mainly initi- ated by solution karst processes. Then, t�anks to t�e tec- tonic downdrop of t�e area, grykes are gradually s�aped by marine processes.

ACKNOwLEDGEMENTS

we are grateful to Fabio C�ersicla for t�e field survey- ing, Dott. Paola Ganis for t�e cluster analysis and Prof.

Franco Stravisi for tide gauge data.

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